As a child, Shiki was taken in as an apprentice to the monster hunter Gurelli. Years later, after Gurelli's death, Shiki returns to his former master's town Akamaya, looking to join a monster hunter's guild. Once in a guild, he meets Irie, whom he discovers is Gurelli's daughter. Shiki partners up with Irie to take up Gurelli's life long goal of hunting down the legendary monster Miogaruna.